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Krebs won the medal in 1953 when the Nobel Prize for Physiology or Medicine was divided equally between Krebs, for his discovery of the citric acid cycle, and Fritz Lipmann, for work on enzymes.

The genes encode enzymes that act in an energy-producing metabolic pathway called the citric acid cycle.

The different profiles of proteins and metabolites clustered into the following groups: fatty acid transport and β-oxidation, gluconeogenesis, and the citric acid cycle.