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An animal rights group argued Missy, Kimba, Lucky, LouLou and Jambo were effectively imprisoned at the zoo, and had filed to have them moved to an elephant sanctuary.

"As with other social justice movements, early losses are expected as we challenge an entrenched status quo that has allowed Missy, Kimba, Lucky, LouLou, and Jambo to be relegated to a lifetime of mental and physical suffering," the group said in a statement.

Coco is 17 years older than the next oldest Twycross chimp Jambo, 41, and 42 years older than it youngest 16-year-old Tuli.

On the morning of the shooting, students sang “Amazing Grace” in the chapel and practiced saying “jambo” — a traditional Swahili greeting — with a missionary doctor who was visiting the school.

Later, when an initial death toll came out, she posted an update: “3 children are confirmed dead. 3 children who learned the word ‘Jambo’ this morning from us in chapel,” she wrote, using the Swahili greeting.