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Four UK shorebirds - the grey plover, dunlin, turnstone and curlew sandpiper - are becoming more endangered on the red list.

Alongside the marine animals, Bischoff said he was excited to find an entire shore ecology that included skulls of sandpipers and pieces of driftwood in the bone bed.

But amid the rivalries, fraud and plagiarism, “The Birds in America,” Audubon’s seminal collection of 435 life-size prints, missed many winged creatures that were not discovered for years, including some common songbirds, hawks and sandpipers.

In spring, two dozen species of shorebirds use the refuge as a way station — primarily Western sandpipers and dunlin.

Without golf balls whizzing overhead, the land has become habitat for migratory shorebirds, among them black-necked stilts, greater yellowlegs and sandpipers, and has even drawn the secretive American bittern.