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Plant-body pseudo-foliaceous with succubous leaf-like lobes.

Stems thalloid, with large subquadrate succubous leaves; underleaves none.

Leaves rather narrow, complicate-bilobed, the lobes subequal or the upper smaller, the lower succubous; underleaves none.

Leaves succubous, bidentate; underleaves 2-cleft, with linear divisions.

Leaves succubous, dorsally decurrent, obliquely ovate-oblong, broadly truncate or bidentate; underleaves smaller, more or less quadrate, bifid or with 4–8 capillary lobes.