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The teleutospore stage of many Uredineae also occurs in deep brown spots.

The teleutospore stages of species of Puccinia, Phragmidium, etc., are also so deep in colour as to appear almost black.

The teleutospore puts forth on germination a four-celled structure, the promycelium or basidium, and this bears later four sporidia or basidiospores, one on each cell.

The teleutospore, with the sporidia which arise from it, is always present, and the division into genera is based chiefly on its characters.

There is also a further reduction in that the basidium is not derived from a teleutospore but is borne directly on the mycelium.